List Of Subgroups Of The Order Coleoptera
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This article classifies the subgroups of the order Coleoptera (
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s) down to the level of
families Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideall ...
, following the system in "Family-group names in Coleoptera (Insecta)", Bouchard, et al. (2011), with corrections and additions from 2020, with common names from bugguide.net. *Order
Coleoptera Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
**Suborder †
Protocoleoptera The Protocoleoptera are a paraphyletic group of extinct beetles, containing the earliest and most primitive lineages of beetles. They represented the dominant group of beetles during the Permian, but were largely replaced by modern beetle groups ...
***Superfamily † Tshekardocoleoidea Rohdendorf, 1944 ****Family † Tshekardocoleidae Rohdendorf, 1944 ****Family † Labradorocoleidae Ponomarenko, 1969 ****Family † Oborocoleidae Kukalová, 1969 ***Superfamily † Permocupedoidea Martynov, 1933 ****Family † Permocupedidae Martynov, 1933 ****Family † Taldycupedidae Rohdendorf, 1961 ***Superfamily † Permosynoidea Tillyard, 1924 ****Family † Ademosynidae Ponomarenko, 1968 ****Family † Permosynidae Tillyard, 1924 **Suborder Archostemata ***Superfamily Cupedoidea Laporte, 1836 ****Family Crowsoniellidae Iablokoff-Khnzorian, 1983 ****Family
Cupedidae The Cupedidae are a small family of beetles, notable for the square pattern of "windows" on their elytra (hard forewings), which give the family their common name of reticulated beetles. The family consists of about 30 species in 9 genera, with ...
Laporte, 1836 ****Family Micromalthidae Barber, 1913 (telephone-pole beetle) ****Family
Ommatidae The Ommatidae are a family of beetles in the suborder Archostemata. The Ommatidae are considered the extant beetle family that has most ancestral characteristics. There are only seven extant species, confined to Australia and South America. How ...
Sharp and Muir, 1912 ****Family Jurodidae Ponomarenko, 1985 ****Family † Triadocupedidae Ponomarenko, 1966 ****Family †
Magnocoleidae Magnocoleidae is an extinct family of beetles in the suborder Archostemata. It was first described in 1998 by Hong, who attributes a single genus, ''Magnocoleus'', to this family. The genus contains a single species ''Magnocoleus huangjiapuensi ...
Hong, 1998 ****Family † Obrieniidae Zherikhin and Gratshev, 1994 ***Superfamily † Asiocoleoidea Rohdendorf, 1961 ****Family † Asiocoleidae Rohdendorf, 1961 ****Family † Tricoleidae Ponomarenko, 1969 ***Superfamily † Rhombocoleoidea Rohdendorf, 1961 ****Family † Rhombocoleidae Rohdendorf, 1961 ***Superfamily † Schizocoleoidea Rohdendorf, 1961 (formerly Schizophoroidea Ponomarenko, 1968) ****Family † Phoroschizidae Bouchard and Bousquet, 2020 (formerly Schizophoridae Ponomarenko, 1968) ****Family † Catiniidae Ponomarenko, 1968 ****Family † Schizocoleidae Rohdendorf, 1961 **Suborder
Myxophaga Myxophaga is the second-smallest suborder of the Coleoptera after Archostemata, consisting of roughly 65 species of small to minute beetles in four families. The members of this suborder are aquatic and semiaquatic, and feed on algae. Descrip ...
***Superfamily Lepiceroidea Hinton, 1936 (1882) ****Family
Lepiceridae ''Lepicerus'' is a genus of myxophagan beetles containing three described species in the family Lepiceridae; it is the only extant genus in the family, with another genus, '' Lepiceratus'' only known from fossils.Jałoszyński, Paweł; Luo, Xiao- ...
Hinton, 1936 (1882) ***Superfamily Sphaeriusoidea Erichson, 1845 ****Family
Torridincolidae The Torridincolidae are a small family of beetles in the suborder Myxophaga. It contains these genera:Hydroscaphidae The Hydroscaphidae are a small family of water beetles known commonly as skiff beetles. As of 2010, there are 23 species in the family. Several are recently described. These beetles are small, most under in length. They are tan to brown in col ...
LeConte, 1874 ****Family
Sphaeriusidae ''Sphaerius'' is a genus of beetles, comprising 23 species, which are the only living members of the family Sphaeriusidae. They are typically found along the edges of streams and rivers, where they feed on algae; they occur on all continents ex ...
Erichson, 1845 **Suborder
Adephaga The Adephaga (from Greek ἀδηφάγος, ''adephagos'', "gluttonous") are a suborder of beetles, and with more than 40,000 recorded species in 10 families, the second-largest of the four beetle suborders. Members of this suborder are collect ...
****Family † Tritarsusidae Hong, 2002 (formerly Tritarsidae Hong, 2002) ****Family
Gyrinidae The whirligig beetles are water beetles, comprising the family Gyrinidae that usually swim on the surface of the water if undisturbed, though they swim underwater when threatened. They get their common name from their habit of swimming rapidly i ...
Latreille, 1810 (whirligig beetles) ****Family
Trachypachidae The Trachypachidae (sometimes known as false ground beetles) are a family of beetles that generally resemble small ground beetles, but that are distinguished by the large coxae of their rearmost legs. There are only six known extant species in th ...
Thomson, 1857 (false ground beetles) ****Family
Rhysodidae Rhysodinae is a subfamily (sometimes called wrinkled bark beetles) in the family Carabidae. There are 19 genera and at least 380 described species in Rhysodinae. The group of genera making up Rhysodinae had been treated as the family Rhysodidae i ...
Laporte, 1840 ****Family
Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it is one of the 10 most species-rich animal fami ...
Latreille, 1802 (ground beetles) ****Family
Haliplidae The Haliplidae are a family of water beetles that swim using an alternating motion of the legs. They are therefore clumsy in water (compared e.g. with the Dytiscidae or Hydrophilidae), and prefer to get around by crawling. The family consists o ...
Aubé, 1836 (crawling water beetles) ****Family † Triaplidae Ponomarenko, 1977 ****Family † Colymbotethidae Ponomarenko, 1994 ****Family †
Parahygrobiidae ''Parahygrobia natans'' is an extinct species of adephagan beetle. It is the only member of the genus ''Parahygrobia'' and family Parahygrobiidae. It is known only from larvae from the Late Jurassic of Russia. Cladistic analysis has placed it a ...
Ponomarenko, 1977 ****Family †
Coptoclavidae Coptoclavidae is an extinct family of aquatic beetles in the suborder Adephaga. The Coptoclavidae lived from the Late Triassic to the Early Cretaceous. Coptoclavidae is a member of the adephagan clade Dytiscoidea, which contains other aquatic b ...
Ponomarenko, 1961 ****Family † Liadytidae Ponomarenko, 1977 ****Family Meruidae Spangler and Steiner, 2005 ****Family
Noteridae Noteridae is a family of adephagan water beetles closely related to the Dytiscidae, and formerly classified with them. They are mainly distinguished by the presence of a distinctive "noterid platform" underneath, in the form of a plate between t ...
Thomson, 1860 (burrowing water beetles) ****Family
Amphizoidae ''Amphizoa'' is a genus of aquatic beetles in the suborder Adephaga, placed in its own monogeneric family, Amphizoidae.Aspidytidae Aspidytidae is a family of aquatic beetles of the suborder Adephaga, described in 2002 from specimens in South Africa and China. There are only two known species in the family and these were originally described in the genus '' Aspidytes,''Beutel ...
Ribera, Beutel, Balke and Vogler, 2002 ****Family
Hygrobiidae ''Hygrobia'' is a genus of aquatic beetles native to Europe, North Africa, China and Australia. It is the only genus in the family Hygrobiidae, also known as the Paelobiidae.Nilsson, A. N. (2006)Which name is valid – Hygrobiidae or Paelobiidae ...
Régimbart, 1879 (1837) ****Family
Dytiscidae The Dytiscidae – based on the Greek ''dytikos'' (δυτικός), "able to dive" – are the predaceous diving beetles, a family of water beetles. They occur in virtually any freshwater habitat around the world, but a few species live a ...
Leach, 1815 (predaceous diving beetles) **Suborder
Polyphaga Polyphaga is the largest and most diverse suborder of beetles. It comprises 144 families in 16 superfamilies, and displays an enormous variety of specialization and adaptation, with over 350,000 described species, or approximately 90% of the b ...
***Infraorder Staphyliniformia ****Superfamily
Histeroidea Histeroidea is a superfamily of beetles in the infraorder Staphyliniformia. Characteristics Characteristic to Histeroidea are an accessory posterior ridge (locking device) behind the hind margin and presence of medial loop and apical hinge of ...
Gyllenhal, 1808 *****Family
Histeridae Histeridae is a family of beetles commonly known as clown beetles or Hister beetles. This very diverse group of beetles contains 3,900 species found worldwide. They can be easily identified by their shortened elytra that leaves two of the seven t ...
Gyllenhal, 1808 *****Family
Sphaeritidae ''Sphaerites'' is a genus of beetles, the only genus in the family Sphaeritidae, sometimes called the false clown beetles. There are five known species, which are widespread in temperate areas of the Northern Hemisphere, found in forested or upla ...
Shuckard, 1839 *****Family
Synteliidae ''Syntelia'' is a genus of beetles. It is the only genus in the family Synteliidae. There are seven known species, which are native to high-elevation regions in southern North America from central Mexico to Guatemala, and in eastern Asia, from ...
Lewis, 1882 ****Superfamily
Hydrophiloidea Hydrophiloidea, known as water scavenger beetles, is a superfamily of beetles. Until recently it included only a single family, the Hydrophilidae (water scavenger beetles), but several of the subfamilies have been removed and raised to family ra ...
Latreille, 1802 *****Family
Epimetopidae Epimetopidae is a family of semi-aquatic beetles belonging to the Hydrophiloidea. They are found in sand and gravel at the edges of streams, rivers and shallow freshwater ponds. These beetles are shorter than half a centimeter long and have a pro ...
Zaitzev, 1908 *****Family
Georissidae ''Georissus'', also called minute mud-loving beetles, is the only genus in the beetle family Georissidae (or Georyssidae). They are tiny insects living in wet soil, often near water. They are found on every continent except Antarctica. Character ...
Laporte, 1840 *****Family Helophoridae Leach, 1815 *****Family Hydrochidae Thomson, 1859 *****Family
Hydrophilidae Hydrophilidae, also known colloquially as water scavenger beetles, is a family of beetles. Aquatic hydrophilids are notable for their long maxillary palps, which are longer than their antennae. Several of the former subfamilies of Hydrophilidae ...
Latreille, 1802 (water scavenger beetles) *****Family Spercheidae Erichson, 1837 ****Superfamily
Staphylinoidea Staphylinoidea is a superfamily of beetles. It is a very large and diverse group with worldwide distribution. Description Adult staphylinoids are generally small beetles no more than a few millimetres long, though Staphylinidae can reach 50 m ...
Latreille, 1802 *****Family
Hydraenidae Hydraenidae is a family of very small aquatic beetles, sometimes called "Minute moss beetles", with a worldwide distribution. They are around 0.8 to 3.3 mm in length. The adults store air on the underside of the body as well as beneath the el ...
Mulsant, 1844 *****Family
Ptiliidae Ptiliidae is a family of very tiny beetles with a cosmopolitan distribution. This family contains the smallest of all beetles, with a length when fully grown of . The weight is approximately 0.4 milligrams. They are colloquially called feat ...
Erichson, 1845 *****Family
Agyrtidae Agyrtidae, or primitive carrion beetles, are a small family of beetles belonging to Staphylinoidea. They are found in mostly temperate areas of the Northern Hemisphere and in New Zealand. Characteristics Agyrtidae are small or middle-sized bee ...
Thomson, 1859 *****Family
Leiodidae Leiodidae is a family of beetles with around 3800 described species found worldwide. Members of this family are commonly called round fungus beetles due to the globular shape of many species, although some are more elongated in shape. They are g ...
Fleming, 1821 *****Family
Silphidae Silphidae is a family of beetles that are known commonly as large carrion beetles, carrion beetles or burying beetles. There are two subfamilies: Silphinae and Nicrophorinae. Nicrophorines are sometimes known as sexton beetles. The number of sp ...
Latreille, 1806 *****Family
Staphylinidae The rove beetles are a family (Staphylinidae) of beetles, primarily distinguished by their short elytra (wing covers) that typically leave more than half of their abdominal segments exposed. With roughly 63,000 species in thousands of genera, the ...
Latreille, 1802 (rove beetles) ***Infraorder
Scarabaeiformia Scarabaeoidea is a superfamily of beetles, the only subgroup of the infraorder Scarabaeiformia. Around 35,000 species are placed in this superfamily and some 200 new species are described each year. Its constituent families are also undergoing ...
****Superfamily
Scarabaeoidea Scarabaeoidea is a superfamily of beetles, the only subgroup of the infraorder Scarabaeiformia. Around 35,000 species are placed in this superfamily and some 200 new species are described each year. Its constituent families are also undergoing r ...
Latreille, 1802 *****Family Pleocomidae LeConte, 1861 (rain beetles) *****Family
Geotrupidae Geotrupidae (from Greek γῆ ''(gē)'', earth, and τρῡπητής ''(trȳpētēs)'', borer) is a family of beetles in the order Coleoptera. They are commonly called earth-boring dung beetles or dor beetles. Most excavate burrows in which ...
Latreille, 1802 (earth-boring scarab beetles) *****Family
Belohinidae ''Belohina inexpectata'' is a species of polyphagan beetles and the sole member of family Belohinidae. It is endemic to southern Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Ma ...
Paulian, 1959 *****Family
Passalidae Passalidae is a family of beetles known variously as "bessbugs", "bess beetles", "betsy beetles" or "horned passalus beetles". Nearly all of the 500-odd species are tropical; species found in North America are notable for their size, ranging fro ...
Leach, 1815 (bess beetles) *****Family
Trogidae Trogidae, sometimes called hide beetles, is a family of beetles with a distinctive warty or bumpy appearance. Found worldwide, the family includes about 300 species contained in four or five genera. Trogids range in length from 2 to 20 mm. Thei ...
MacLeay, 1819 (hide beetles) *****Family
Glaresidae ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy area ...
Prudhomme de Borre, 1886 *****Family Diphyllostomatidae Holloway, 1972 *****Family
Lucanidae Stag beetles are a family of about 1,200 species of beetles in the family Lucanidae, currently classified in four subfamilies.Smith, A.B.T. (2006). A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections ...
Latreille, 1804 (stag beetles) *****Family
Ochodaeidae Ochodaeidae, also known as the sand-loving scarab beetles, is a small family of scarabaeiform beetles occurring in many parts of the world. These beetles are small, ranging from . Their bodies are elongate and convex, with black and brown colo ...
Streubel, 1846 (sand-loving scarab beetles) *****Family
Hybosoridae Hybosoridae, sometimes known as the scavenger scarab beetles, is a family of scarabaeiform beetles. The 690 species in 97 genera occur widely in the tropics, but little is known of their biology. Hybosorids are small, 5–7 mm in length a ...
Erichson, 1847 *****Family
Glaphyridae Glaphyridae is a family of beetles, commonly known as bumble bee scarab beetles. There are eight extant genera with about 80 species distributed worldwide and two extinct genera described from the Aptian aged Yixian Formation of China. There are ...
MacLeay, 1819 (bumble bee scarab beetles) *****Family
Scarabaeidae The family Scarabaeidae, as currently defined, consists of over 30,000 species of beetles worldwide; they are often called scarabs or scarab beetles. The classification of this family has undergone significant change in recent years. Several sub ...
Latreille, 1802 (scarab beetles) *****Family † Coprinisphaeridae Genise, 2004 (
ichnotaxon An ichnotaxon (plural ichnotaxa) is "a taxon based on the fossilized work of an organism", i.e. the non-human equivalent of an artifact. ''Ichnotaxa'' comes from the Greek ίχνος, ''ichnos'' meaning ''track'' and ταξις, ''taxis'' meaning ...
) *****Family † Pallichnidae Genise, 2004 (ichnotaxon) ***Infraorder
Elateriformia Elateriformia is an infraorder of polyphagan beetles. The two largest families in this group are buprestids, of which there are around 15,000 described species, and click beetles, of which there are around 10,000 described species. The infraor ...
****Superfamily
Scirtoidea Scirtoidea is a superfamily of beetles. It is traditionally considered to consist of four families: Clambidae, Decliniidae, Eucinetidae and Scirtidae. However, genetic studies have suggested that Clambidae and Eucinetidae belong to a separate s ...
Fleming, 1821 *****Family
Decliniidae Decliniidae is a family of beetles belonging to Scirtoidea. It contains the single genus ''Declinia'' with two species, ''D. relicta'' and ''D. versicolor'', found in the Russian Far East and Japan, respectively. Little is known of their ecology, ...
Nikitsky, Lawrence, Kirejtshuk and Gratshev, 1994 *****Family
Eucinetidae Eucinetidae is a family of beetles, notable for their large coxal plates that cover much of the first ventrite of the abdomen, sometimes called plate-thigh beetles. The family is small for beetles, with about 50 species in 11 genera, but are ...
Lacordaire, 1857 *****Family
Clambidae Clambidae is a family of beetles. They are known commonly as the minute beetlesMajka, C. G., & Langor, D. (2009)Clambidae (Coleoptera) of Atlantic Canada.''Journal of the Acadian Entomological Society'' 5(7), 32-40. or the fringe-winged beetles.< ...
Fischer von Waldheim, 1821 *****Family
Scirtidae Scirtidae is a Taxonomic rank, family of beetles (Coleoptera). These beetles are commonly referred to as marsh beetles, as the larvae are typically associated with stagnant water, but can be found in flowing water. Adults prefer decomposing plan ...
Fleming, 1821 *****Family † Elodophthalmidae Kirejtshuk and Azar, 2008 *****Family † Mesocinetidae Kirejtshuk and Ponomarenko, 2010 ****Superfamily Dascilloidea Guérin-Méneville, 1843 (1834) *****Family
Dascillidae Dascillidae is a family of beetles within the clade Elateriformia. There are about 100 extant species in 11 genera, which are found worldwide. Dascillidae together with Rhipiceridae form the super family Dascilloidea. Taxonomy The family was na ...
Guérin-Méneville, 1843 (1834) *****Family
Rhipiceridae Rhipiceridae is a family of beetles found worldwide. The larva of rhipicerids are parasitoids of cicada nymphs. Rhipiceridae and Dascillidae form the super family Dascilloidea, within the Elateriformia. Taxonomic History The taxonomic history ...
Latreille, 1834 ****Superfamily
Buprestoidea Buprestoidea is a superfamily of beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insect ...
Leach, 1815 *****Family Schizopodidae LeConte, 1859 *****Family
Buprestidae Buprestidae is a family of beetles known as jewel beetles or metallic wood-boring beetles because of their glossy iridescent colors. Larvae of this family are known as flatheaded borers. The family is among the largest of the beetles, with some ...
Leach, 1815 ****Superfamily Byrrhoidea Latreille, 1804 *****Family Byrrhidae Latreille, 1804 *****Family Protelmidae Jeannel, 1950 *****Family
Elmidae Elmidae, commonly known as riffle beetles, is a family (biology), family of beetles in the superfamily Byrrhoidea described by John Curtis (entomologist), John Curtis in 1830. Both adults and larvae are usually aquatic, living under rocks in fas ...
Curtis, 1830 *****Family
Dryopidae Dryopidae is a family of beetles, commonly named long-toed water beetles, in the superfamily Byrrhoidea. It was described by Gustaf Johan Billberg in 1820. Description Long-toed water beetles are named for their extended claws. Adults have dense ...
Billberg, 1820 (1817) *****Family
Lutrochidae Lutrochidae is a family of water beetles with a single genus ''Lutrochus'' sometimes known as "travertine beetles". There are around 21 species native to the Americas from the southern United States to Brazil. They are distinguished by their ov ...
Kasap and Crowson, 1975 *****Family
Limnichidae Limnichidae, commonly called minute marsh-loving beetles, is a family of beetles belonging to Byrrhoidea. There are at least 30 genera and 350 described species in Limnichidae. They are found worldwide, with the greatest diversity in tropical regi ...
Erichson, 1846 *****Family
Heteroceridae Heteroceridae, the variegated mud-loving beetles, are a widespread and relatively common family of beetles found on every continent except for Antarctica. Around two hundred and fifty species of heterocerids are known to occur worldwide. They a ...
MacLeay, 1825 *****Family
Psephenidae Water-penny beetles are a family (the Psephenidae) of 272 species (in 35 genera) of aquatic beetles found on all continents except Antarctica, in both tropical and temperate areas. The young, which live in water, resemble tiny pennies. The larv ...
Lacordaire, 1854 *****Family
Cneoglossidae Cneoglossidae is a family of beetles in the superfamily Byrrhoidea, containing nine described species in a single genus, ''Cneoglossa,'' which are native to the Neotropics The Neotropical realm is one of the eight biogeographic realms const ...
Champion, 1897 *****Family
Ptilodactylidae Ptilodactylidae is a family of beetles belonging to the Elateriformia. There around 500 extant species in 35 genera. They are generally associated with riparian and aquatic habitats. The larvae generally live associated with rotting wood or veg ...
Laporte, 1836 *****Family Podabrocephalidae Pic, 1930 *****Family Chelonariidae Blanchard, 1845 *****Family
Eulichadidae Eulichadidae is a family of beetles belonging to Elateriformia. There are two extant genera, '' Eulichas'' with several dozen species native to the Indomalayan realm of Asia, and '' Stenocolus,'' with a single species native to Western North Ame ...
Crowson, 1973 *****Family Callirhipidae Emden, 1924 ****Superfamily
Elateroidea The Elateroidea are a large superfamily of beetles. It contains the familiar click beetles, fireflies, and soldier beetles and their relatives. It consists of about 25,000 species. Description Elateroidea is a morphologically diverse group, i ...
Leach, 1815 *****Family
Artematopodidae Artematopodidae is a family of soft-bodied plant beetles in the superfamily Elateroidea. They are mostly found in understory forest foliage. The life history of the group is obscure, larvae of the genera ''Eurypogon'' and ''Macropogon'' likely fee ...
Lacordaire, 1857 *****Family
Brachypsectridae The Brachypsectridae are a family of beetles commonly known as the Texas beetles. There are only two extant genera, '' Brachypsectra'' and '' Asiopsectra. Brachypsectra'' has a cosmopolitan distribution, mostly in arid regions, while ''Asiopsectr ...
Horn, 1881 *****Family
Cantharidae The soldier beetles (Cantharidae) are relatively soft-bodied, straight-sided beetles. They are cosmopolitan in distribution. One of the first described species has a color pattern reminiscent of the Red coat (British army), red coats of early Bri ...
Imhoff, 1856 (1815) (soldier beetles) *****Family
Cerophytidae The Cerophytidae are a family of beetles belonging to Elateroidea. Larvae are associated with rotting wood, on which they are presumed to feed.Costa, Cleide, Vanin, Sergio A., Lawrence, John F. and Ide, Sergio. "4.4. Cerophytidae Latreille, 1834" ...
Latreille, 1834 *****Family
Elateridae Elateridae or click beetles (or "typical click beetles" to distinguish them from the related families Cerophytidae and Eucnemidae, which are also capable of clicking) are a family of beetles. Other names include elaters, snapping beetles, spr ...
Leach, 1815 (click beetles; includes former Drilidae, Omalisidae, and Plastoceridae) *****Family
Eucnemidae Eucnemidae, or false click beetles, are a family of elateroid beetles including about 1700 species distributed worldwide. Description Closely related to the family Elateridae, specimens of Eucnemidae can reach a length of . Bodies are slightl ...
Eschscholtz, 1829 *****Family
Lampyridae The Lampyridae are a family (biology), family of Elateroidea, elateroid beetles with more than 2,000 described species, many of which are bioluminescence, light-emitting. They are soft-bodied beetles commonly called fireflies, lightning bugs, ...
Rafinesque, 1815 (fireflies) *****Family
Lycidae The Lycidae are a family in the beetle order Coleoptera, members of which are commonly called net-winged beetles. These beetles are cosmopolitan, being found in Nearctic, Palearctic, Neotropical, Afrotropical, Oriental, and Australian ecoregions ...
Laporte, 1836 *****Family
Omethidae Omethidae is a family of Elateroidea sometimes known as the false soldier beetles. They are native to South, Southeast and Eastern Asia and the Americas. Their biology is obscure and their larvae are unknown. They appear to inhabit vegetation in ...
LeConte, 1861 (includes former Telegeusidae) *****Family
Phengodidae The beetle family Phengodidae is known also as glowworm beetles, whose larvae are known as glowworms. The females and larvae have bioluminescence, bioluminescent organs. They occur throughout the New World from extreme southern Canada to Chile. ...
LeConte, 1861 *****Family
Rhagophthalmidae The Rhagophthalmidae are a family of beetles within the superfamily Elateroidea. Members of this beetle family have bioluminescent organs on the larvae, and sometimes adults, and are closely related to the Phengodidae (glowworm beetles), though ...
Olivier, 1907 *****Family Sinopyrophoridae Bi, 2018 *****Family
Throscidae Throscidae is a family of elateroid beetles found worldwide (except New Zealand) with around 150 species in 5 extant genera. The larvae are soil-dwelling, siphoning fluid from mycorrhizae attached to trees. The adults are short-lived, with the ...
Laporte, 1840 nomen protectum *****Family † Berendtimiridae Winkler, 1987 *****Family † Praelateriidae Dolin, 1973 ****Superfamily Rhinorhipoidea Lawrence, 1988 *****Family Rhinorhipidae Lawrence, 1988 ***Infraorder
Bostrichiformia Bostrichiformia is an infraorder of polyphagan beetles. It contains two superfamilies, Derodontoidea and Bostrichoidea, which includes the Dermestidae, Ptinidae Ptinidae is a family of beetles in the superfamily Bostrichoidea. There are ...
****Superfamily Derodontoidea LeConte, 1861 *****Family
Derodontidae Derodontidae is a family of beetles, in its own superfamily, Derodontoidea, sometimes known as the tooth-necked fungus beetles. Beetles of this family are small, between 2 and 6 mm in length, typically with spiny margins on their pronotum (p ...
LeConte, 1861 *****Family
Nosodendridae Nosodendridae is a family of beetles, with only 67 species in three extant genera, which are found worldwide. '' Nosodendron'', the largest genus, is found in forests and attracted to yeast generated slime on the wounds of trees, and likely consu ...
Erichson, 1846 *****Family
Jacobsoniidae Jacobsoniidae are a family of tiny beetles belonging to Staphylinoidea. The larvae and adults live under bark, in plant litter, fungi, bat guano and rotten wood. There are around 28 described species in three genera: Description Members of this ...
Heller, 1926 ****Superfamily Bostrichoidea Latreille, 1802 *****Family
Dermestidae Dermestidae are a family of Coleoptera that are commonly referred to as skin beetles. Other common names include larder beetle, hide or leather beetles, carpet beetles, and khapra beetles. There are over 1,100 species described. Dermestids have ...
Latreille, 1804 (carpet beetles) *****Family Endecatomidae LeConte, 1861 *****Family
Bostrichidae The Bostrichidae are a family of beetles with more than 700 described species. They are commonly called auger beetles, false powderpost beetles, or horned powderpost beetles. The head of most auger beetles cannot be seen from above, as it is down ...
Latreille, 1802 *****Family
Ptinidae Ptinidae is a family of beetles in the superfamily Bostrichoidea. There are at least 220 genera and 2,200 described species in Ptinidae worldwide. The family includes spider beetles and deathwatch beetles. The Ptinidae family species are hard ...
Latreille, 1802 ***Infraorder
Cucujiformia Cucujiformia is an infraorder of polyphagan beetles, representing most plant-eating beetles. The infraorder contains the seven superfamilies: * Chrysomeloidea (~7 families including longhorn beetles and leaf beetles) * Cleroidea (checkered bee ...
****Superfamily Lymexyloidea Fleming, 1821 *****Family
Lymexylidae The Lymexylidae (historically often spelled Lymexylonidae), also known as ship-timber beetles, are a family of wood-boring beetles. Lymexylidae belong to the suborder Polyphaga and are the sole member of the superfamily Lymexyloidea. Habitat a ...
Fleming, 1821 ****Superfamily
Cleroidea Cleroidea is a small superfamily of beetles containing over 10,000 species. Most of the members of the group are somewhat slender, often with fairly soft, flexible elytra, and typically hairy or scaly. Description Cleroidea is defined by the fo ...
Latreille, 1802 *****Family
Phloiophilidae ''Phloiophilus edwardsii'' is the sole known species of the beetle family Phloiophilidae in the superfamily Cleroidea. It is native to Europe. The larvae are mycophagous, and have been observed feeding on basidiomycetes of the genus ''Phlebia ...
Kiesenwetter, 1863 *****Family
Trogossitidae Trogossitidae, also known as bark-gnawing beetles, are a small family in the superfamily Cleroidea. Many taxa formerly within this family have been removed (as of 2019) to other families, such as Lophocateridae, Peltidae, Protopeltidae, Rentoni ...
Latreille, 1802 *****Family
Chaetosomatidae Chaetosomatidae is a small family of beetles, in the superfamily Cleroidea Cleroidea is a small superfamily of beetles containing over 10,000 species. Most of the members of the group are somewhat slender, often with fairly soft, flexible elyt ...
Crowson, 1952 *****Family Metaxinidae Kolibáč, 2004 *****Family
Thanerocleridae Thanerocleridae is a family of beetles belonging to the superfamily Cleroidea. It was formerly considered a subfamily of Cleridae, but was recently elevated to the rank of family. The family has 36 living species in 10 genera, which are found glo ...
Chapin, 1924 *****Family
Cleridae Cleridae are a family of beetles of the superfamily Cleroidea. They are commonly known as checkered beetles. The family Cleridae has a worldwide distribution, and a variety of habitats and feeding preferences. Cleridae have many niches and fe ...
Latreille, 1802 (checkered beetles) *****Family Acanthocnemidae Crowson, 1964 *****Family
Phycosecidae Phycosecidae is a family of beetles in the superfamily Cleroidea Cleroidea is a small superfamily of beetles containing over 10,000 species. Most of the members of the group are somewhat slender, often with fairly soft, flexible elytra, and typ ...
Crowson, 1952 *****Family Prionoceridae Lacordaire, 1857 *****Family Mauroniscidae Majer, 1995 *****Family
Melyridae Melyridae (common name: soft-winged flower beetles) are a family of beetles of the superfamily Cleroidea. Description Most are elongate-oval, soft-bodied beetles 10 mm long or less. Many are brightly patterned in black and brown, yellow, ...
Leach, 1815 (soft-winged flower beetles) ****Superfamily
Cucujoidea Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark ...
Latreille, 1802 *****Family † Parandrexidae Kirejtshuk, 1994 *****Family † Sinisilvanidae Hong, 2002 *****Family
Boganiidae Boganiidae is a family of beetles, in the superfamily Cucujoidea. Members of the family are found in southern Africa, Australia and New Caledonia. Adults and larvae are pollenivorous, feeding on the pollen of cycads and flowering plants of the f ...
Sen Gupta and Crowson, 1966 *****Family
Byturidae Byturidae, also known as Fruitworms, is a very small family of beetles, in the suborder Polyphaga, comprising fewer than 20 species worldwide.C.A. Springer & M.A. Goodrich (1994) A Revision of the Subfamily Platydascillinae (Coleoptera: Byturidae ...
Gistel, 1848 *****Family
Helotidae Helotidae is a family of beetles, in the suborder Polyphaga. The family includes about five extant genera, ''Helota'' MacLeay, ''Neohelota'' Ohta, ''Afrohelotina'' Kirejtshuk, ''Metahelotella'' Kirejtshuk, and ''Strophohelota'' Kirejtshuk. Hel ...
Chapuis, 1876 *****Family Protocucujidae Crowson, 1954 *****Family Sphindidae Jacquelin du Val, 1860 *****Family
Biphyllidae Biphyllidae, or false skin beetles, are a family of beetles, in the superfamily Cleroidea. The have a cosmopolitan distribution (excluding New Zealand). About 195 species are known. They live under the bark of dead trees and in leaf litter, and a ...
LeConte, 1861 *****Family
Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present circumscription, it contains 6 tribes (Tritomini, Dacnini, Megalodacnini, Er ...
Latreille, 1802 *****Family
Monotomidae Monotomidae is a family of beetles in the superfamily Cucujoidea. The family is found worldwide, with approximately 240 species in 33 genera. The ecological habits of the family are diverse, with different members of the group being found under t ...
Laporte, 1840 *****Family
Hobartiidae Hobartiidae is a family of beetles, in the superfamily Cucujoidea. There are only two known genera '' Hobartius'' (Sen Gupta & Crowson, 1966) and '' Hydnobioides'' (Sen Gupta & Crowson, 1966), with six species, five of which are native to Austral ...
Sen Gupta and Crowson, 1966 *****Family
Cryptophagidae Cryptophagidae is a family of beetles with representatives found in all biogeographic realms. Members of this family are commonly called silken fungus beetles and both adults and larvae appear to feed exclusively on fungi although in a wide vari ...
Kirby, 1826 *****Family Agapythidae Sen Gupta and Crowson, 1969 *****Family Priasilphidae Crowson, 1973 *****Family Phloeostichidae Reitter, 1911 *****Family
Silvanidae Silvanidae, "silvan flat bark beetles", is a family of beetles in the superfamily Cucujoidea,Thomas, M. C., and R.A. B. Leschen. 2010. Silvanidae Kirby, 1837. p. 346-350. In: Leschen, R.A.B., R.G. Beutel, and J.F. Lawrence. Coleoptera, Beetles. Vo ...
Kirby, 1837 *****Family
Cucujidae The Cucujidae, "flat bark beetles," are a family of distinctively flat beetles found worldwide (except Africa and Antarctica) under the bark of dead trees. The family has received considerable taxonomic attention in recent years and now consists ...
Latreille, 1802 *****Family
Myraboliidae ''Myrabolia'' is the only genus in the beetle family Myraboliidae in the superfamily Cucujoidea. It has about 13 species, found in Australia. Adults and possibly larvae live under the bark of ''Eucalyptus ''Eucalyptus'' () is a genus of over s ...
Lawrence and Britton, 1991 *****Family
Cavognathidae Cavognathidae is a family of beetles, in the superfamily Cucujoidea Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles and ...
Sen Gupta and Crowson, 1966 *****Family
Lamingtoniidae ''Lamingtonium'' is the only genus in the family Lamingtoniidae, of the beetle superfamily Cucujoidea. It contains three species endemic to Australia.Lawrence, J. F., and Leschen, R. A. B. (2003). Review of Lamingtoniidae (Coleoptera: Cucujoidea ...
Sen Gupta and Crowson, 1969 *****Family Passandridae Blanchard, 1845 *****Family
Phalacridae The Phalacridae are a family of beetles commonly called the shining flower beetles, They are often found in composite flowers. They are oval-shaped, usually tan, and aboout 2 mm in length. Most species feed on fungus, although a number feed ...
Leach, 1815 *****Family Propalticidae Crowson, 1952 *****Family
Laemophloeidae Laemophloeidae, "lined flat bark beetles," is a family in the superfamily Cucujoidea characterized by predominantly dorso-ventrally compressed bodies, head and pronotal discs bordered by ridges or grooves, and inverted male genitalia. Size range ...
Ganglbauer, 1899 *****Family Tasmosalpingidae Lawrence and Britton, 1991 *****Family
Cyclaxyridae Cyclaxyridae are a family of beetles in the superfamily Cucujoidea. The only living genus is ''Cyclaxyra'', with two species endemic to New Zealand. Other species have been named from fossils. They are also known as sooty mould beetles due to the ...
Gimmel, Leschen and Ślipiński, 2009 *****Family
Kateretidae Kateretidae also known as short-winged flower beetles are a family of beetles in the superfamily Cucujoidea. There are 10 extant and 4 extinct genera, and at least 40 described species. They are found worldwide except in New Zealand. Adults are a ...
Kirby, 1837 *****Family
Nitidulidae The sap beetles, also known as Nitidulidae, are a family (biology), family of beetles. They are small (2–6 mm) ovoid, usually dull-coloured beetles, with knobbed antenna (biology), antennae. Some have red or yellow spots or bands. They fe ...
Latreille, 1802 *****Family Smicripidae Horn, 1880 *****Family Bothrideridae Erichson, 1845 *****Family Cerylonidae Billberg, 1820 *****Family Alexiidae Imhoff, 1856 *****Family Discolomatidae Horn, 1878 *****Family
Endomychidae Endomychidae, or handsome fungus beetles, is a family of beetles with representatives found in all biogeographic realms. There are around 120 genera and 1300 species. The family was established based on the type genus '' Endomychus'', a genus ere ...
Leach, 1815 *****Family
Coccinellidae Coccinellidae () is a widespread family of small beetles ranging in size from . They are commonly known as ladybugs in North America and ladybirds in Great Britain. Some entomologists prefer the names ladybird beetles or lady beetles as they ...
Latreille, 1807 (ladybirds or lady beetles) *****Family Corylophidae LeConte, 1852 *****Family Akalyptoischiidae Lord, Hartley, Lawrence, McHugh, Whiting and Miller, 2010 *****Family
Latridiidae Latridiidae (sometimes spelled "Lathridiidae") is a family of tiny, little-known beetles commonly called minute brown scavenger beetles or fungus beetles. The number of described species currently stands at around 1050 in 29 genera but the number ...
Erichson, 1842 ****Superfamily
Tenebrionoidea The Tenebrionoidea are a very large and diverse superfamily of beetles. It generally corresponds to the Heteromera of earlier authors. Taxonomy Tenebrionoidea contains the following families: * Aderidae Winkler 1927 (ant-like leaf beetles) ...
Latreille, 1802 *****Family
Mycetophagidae The Mycetophagidae or hairy fungus beetles are a family of beetles in the superfamily Tenebrionoidea. The different species are between 1.0 and 6.5 mm in length. The larvae and adults live in decaying leaf litter, fungi, and under bark. Mos ...
Leach, 1815 *****Family Archeocrypticidae Kaszab, 1964 *****Family
Pterogeniidae Pterogeniidae is a family of beetles belonging to the supefamily Tenebrionoidea. They are found in South Asia, Southeast Asia, China, New Guinea and the Solomon Islands. Adults and larvae have been found associated with the fruiting bodies of pol ...
Crowson, 1953 *****Family
Ciidae The minute tree-fungus beetles, family (biology), family Ciidae, are a sizeable group of beetles which inhabit Polyporales bracket fungi or coarse woody debris. Most numerous in warmer regions, they are nonetheless widespread and a considerable n ...
Leach, 1819 *****Family
Tetratomidae Tetratomidae is a small family of beetles sometimes called polypore fungus beetles. The family consists of several genera, most of which used to be in the family Melandryidae. Tetratomidae can be found worldwide. Their food consists of fruiti ...
Billberg, 1820 *****Family
Melandryidae Melandryidae is a family of beetles in the superfamily Tenebrionoidea. Members of the family are found worldwide, with around 420 species in 60 genera. Larvae and adults are generally associated with rotting wood and wood-decomposing fungi. Ge ...
Leach, 1815 *****Family
Mordellidae The Mordellidae are a family of beetles commonly known as tumbling flower beetles for the typical irregular movements they make when escaping predators, or as pintail beetles due to their abdominal tip which aids them in performing these tumbling ...
Latreille, 1802 (tumbling flower beetles) *****Family
Ripiphoridae Ripiphoridae (formerly spelled Rhipiphoridae) is a cosmopolitan family of some 450 described species of beetles sometimes called "wedge-shaped beetles". Ripiphoridae are unusual among beetle families in that many species are hypermetamorphic pa ...
Laporte, 1840 *****Family
Zopheridae The Zopheridae family of beetles has grown considerably in recent years as the members of two other families have been included within its circumscription; these former families are the Monommatidae and the Colydiidae, which are now both incl ...
Solier, 1834 *****Family Ulodidae Pascoe, 1869 *****Family
Promecheilidae Promecheilidae is a family of beetles in the superfamily Tenebrionoidea. They were formerly included in a family called Perimylopidae. They are found in southern South America and associated archipelagos like South Georgia and the Falklands, Ne ...
Lacordaire, 1859 *****Family
Chalcodryidae The Chalcodryidae are a family of beetles in the superfamily Tenebrionoidea. It contains at least five species in two genera '' Chalcodrya'' and '' Philpottia,'' which are endemic to New Zealand. They are generally found associated with moss or ...
Watt, 1974 *****Family Trachelostenidae Lacordaire, 1859 *****Family
Tenebrionidae Darkling beetle is the common name for members of the beetle family Tenebrionidae. The number of species in the Tenebrionidae is estimated at more than 20,000 and the family is cosmopolitan in distribution. Taxonomy ''Tenebrio'' is the Latin gen ...
Latreille, 1802 (darkling beetles) *****Family
Prostomidae Prostomidae is a family of beetles with no vernacular common name, though recent authors have coined the name jugular-horned beetles. They are often found in dead wood. The family consist of two extant genera with about 20 species. '' Prostomis a ...
Thomson, 1859 *****Family Synchroidae Lacordaire, 1859 *****Family Stenotrachelidae Thomson, 1859 *****Family
Oedemeridae The family Oedemeridae is a cosmopolitan group of beetles commonly known as false blister beetles, though some recent authors have coined the name pollen-feeding beetles. There are some 100 genera and 1,500 species in the family, mostly associate ...
Latreille, 1810 *****Family
Meloidae Blister beetles are beetles of the family Meloidae, so called for their defensive secretion of a blistering agent, cantharidin. About 7,500 species are known worldwide. Many are conspicuous and some are aposematically colored, announcing their ...
Gyllenhal, 1810 (blister beetles) *****Family
Mycteridae The family Mycteridae is a small group of tenebrionoid beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them f ...
Perty, 1840 (palm beetles and flower beetles) *****Family
Boridae The Boridae are a small family of tenebrionoid beetles with no vernacular common name, though recent authors have coined the name conifer bark beetles. The family contains three genera. '' Boros'' is native to North America and northern Eurasia, ...
Thomson, 1859 *****Family
Trictenotomidae The Trictenotomidae are a small family of beetles in the superfamily Tenebrionoidea, containing fifteen species in two genera. Most species are found in the Oriental realm where they live in montane forest habitats. The family is considered, bas ...
Blanchard, 1845 *****Family
Pythidae The family Pythidae is a small group of tenebrionoid beetles with no vernacular common name, though recent authors have coined the name dead log bark beetles. There are seven genera, which are largely native to the mid-high latitude regions of t ...
Solier, 1834 *****Family Pyrochroidae Latreille, 1806 *****Family
Salpingidae Salpingidae or narrow-waisted bark beetles is a family of beetles in the superfamily Tenebrionoidea. The species are small, about 1.5 – 7 mm in length. The family is globally distributed and consists of about 45 genera and 300 species, w ...
Leach, 1815 *****Family
Anthicidae The Anthicidae are a family of beetles that resemble ants. They are sometimes called ant-like flower beetles or ant-like beetles. The family comprises over 3,500 species in about 100 genera. Description Their heads constrict just in front of ...
Latreille, 1819 *****Family
Aderidae The Aderidae, sometimes called ant-like leaf beetles, are a family of beetles that bear some resemblance to ants. The family consists of about 1,000 species in about 40 genera, of which most are tropical, although overall distribution is worldwid ...
Csiki, 1909 *****Family Scraptiidae Gistel, 1848 ****Clade
Phytophaga Phytophaga is a clade of beetles within the infraorder Cucujiformia consisting of the superfamilies Chrysomeloidea and Curculionoidea that are distinctive in the plant-feeding habit combined with the tarsi being pseudotetramerous or cryptopentam ...
*****Superfamily
Chrysomeloidea The Chrysomeloidea are an enormous superfamily of beetles, with tens of thousands of species. The largest families are Cerambycidae, long-horned beetles, with more than 35,000 species, and Chrysomelidae, leaf beetles, with more than 13,000 specie ...
Latreille, 1802 ******Family
Oxypeltidae The Oxypeltidae are a small family belonging to the superfamily Chrysomeloidea, widespread in the Andean region of Chile and Argentina. They have traditionally been considered a group within the Cerambycidae. Description The body is massive and ...
Lacordaire, 1868 ******Family Vesperidae Mulsant, 1839 ******Family
Disteniidae The Disteniidae are a small family of beetles in the superfamily Chrysomeloidea, traditionally treated as a group within the Cerambycidae (most resembling certain Lepturinae such as the genus '' Stenocorus'', but having long fine antennae and s ...
Thomson, 1861 ******Family
Cerambycidae The longhorn beetles (Cerambycidae), also known as long-horned or longicorns, are a large family of beetles, with over 35,000 species described. Most species are characterized by extremely long antennae, which are often as long as or longer than ...
Latreille, 1802 (longhorn beetles) ******Family
Megalopodidae The Megalopodidae are a small family of leaf beetles, previously included as a subfamily within the Chrysomelidae. One of its constituent subfamilies, Zeugophorinae, has also frequently been treated as a subfamily within Chrysomelidae. The fami ...
Latreille, 1802 ******Family Orsodacnidae Thomson, 1859 ******Family
Chrysomelidae The insects of the beetle family Chrysomelidae are commonly known as leaf beetles, and include over 37,000 (and probably at least 50,000) species in more than 2,500 genera, making up one of the largest and most commonly encountered of all beetle ...
Latreille, 1802 (leaf beetles) *****Superfamily
Curculionoidea Weevils are beetles belonging to the superfamily Curculionoidea, known for their elongated snouts. They are usually small, less than in length, and herbivorous. Approximately 97,000 species of weevils are known. They belong to several families, ...
Latreille, 1802 ******Family
Nemonychidae Nemonychidae is a small family of weevils, placed within the ''primitive weevil'' group because they have straight rather than geniculate (elbowed) antennae. They are often called pine flower weevils. As in the Anthribidae, the labrum appears ...
Bedel, 1882 ******Family
Anthribidae Anthribidae is a family of beetles also known as fungus weevils. The antennae are not elbowed, may occasionally be longer than the body and thread-like, and can be the longest of any members of Curculionoidea. As in the Nemonychidae, the labrum ...
Billberg, 1820 ******Family † Ulyanidae Zherikhin, 1993 ******Family
Belidae Belidae is a family of weevils, called belids or primitive weevils because they have straight antennae, unlike the "true weevils" or Curculionidae which have geniculate (elbowed) antennae. They are sometimes known as "cycad weevils", but this pro ...
Schönherr, 1826 ******Family
Caridae Caridae is a small Gondwanan family of weevils. They are considered part of the primitive weevil group, because they have straight rather than geniculate (elbowed) antennae. The insertion of the antennae on the rostrum cannot be seen from above. ...
Thompson, 1992 ******Family
Attelabidae The Attelabidae is a widespread family of weevils. They are among the primitive weevils, because of their straight antennae, which are inserted near the base of the rostrum. The prothorax is much narrower than the base of the elytra on the abdo ...
Billberg, 1820 ******Family
Brentidae Brentidae is a cosmopolitan family of primarily xylophagous beetles also known as straight-snouted weevils. The concept of this family has been recently expanded with the inclusion of three groups formerly placed in the Curculionidae; the subfa ...
Billberg, 1820 ******Family
Brachyceridae Brachyceridae is a family of weevils. There are at least 150 genera in Brachyceridae. It was treated as a subfamily of Curculionidae. Cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in c ...
Billberg, 1820 ******Family
Curculionidae The Curculionidae are a family of weevils, commonly called snout beetles or true weevils. They are one of the largest animal families, with 6,800 genera and 83,000 species described worldwide. They are the sister group to the family Brentidae. T ...
Latreille, 1802 (snout beetles, weevils, and bark beetles)


References

* Lawrence, J.F., Newton, A.F. Jr. (1995) Families and subfamilies of Coleoptera (with selected genera, notes, references, and data on family-group names), pp. 779–1006. In: Pakaluk, J., Slipinski, S.A. (eds.), ''Biology, phylogeny, and classification of Coleoptera: Papers celebrating the 80th birthday of Roy A. Crowson''. Muzeum i Instytut Zoologii PAN, Warszawa. *
Ross H. Arnett, Jr. Ross Harold Arnett Jr. (April 13, 1919 – July 16, 1999) was an American entomologist noted for his studies of beetles, and as founder of the ''Coleopterist's Bulletin''. Born in Medina, New York, he was a star student at Cornell University, wher ...
and Michael C. Thomas, ''
American Beetles ''American Beetles'' is the single most comprehensive description of the beetles of North America north of the tropical area of Mexico. It was started by Ross H. Arnett, Jr. as an update of his classic ''The Beetles of the United States''; along ...
'' (
CRC Press The CRC Press, LLC is an American publishing group that specializes in producing technical books. Many of their books relate to engineering, science and mathematics. Their scope also includes books on business, forensics and information tec ...
, 2001–2002) {{Coleoptera *